troubleshooting Question

Cannot route internal messages - server to server - Exchange 2003 SP2

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dcheyetteFlag for United States of America asked on
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Exchange 2003 - SP2
Server1 - Original Server (swing method upgrade from Exchange 5.5 about 6 months ago)
Server2 - New Last week (lives over VPN connection from Server 1)

Last week I moved mailboxes from Server1 to Server2 so all users had local mailboxes.  

We also have 2 internet domains - domain1 smtp mail comes to server1 directly from internet.
                                                  domain2 smtp mail comes to server2 directly from internet.

Problem is... when a user from server1 tries to email a user whose mailbox store is on server2, the mail sits in the queue (Labeled Default SMTP Virtual Server).

Additionally, Server2 users can SUCCESSFULLY send mail to users on Server1, so this is a one-way problem.

Other information:

- Server2 users have a default internet email address of @domain2.com
- Recipient Policies are set to allow both domains.

I don't understand why this should be an SMTP issue as these mailboxes should be found by server1 internally.  Active Directory U&C shows proper mail store.  Replication between DC's is working properly.

Yes, I can ping servers in both directions.

Ideas?
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