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Exchange is the server side of a collaborative application product that is part of the Microsoft Server infrastructure. Exchange's major features include email, calendaring, contacts and tasks, support for mobile and web-based access to information, and support for data storage.
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You have one Mailbox store per Storage Group! You could have just left them all in one for easier management. Each Storage Group can have up to 5 stores, so with your 4 stores and the public, that would be maxing out one storage group.