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Exchange 2003 to 2007 mail in 'unreachable destination' queue

Last Modified: 2013-11-15
1. Windows Server 2003 R2 Standard SP2 w/ Exchange 2003 SP2
2. Windows Server 2008 Standard x64 SP2 w/ Exchange 2007 SP2

- Installation logs for Exchange 2007 show successful installation.

- Can send mail from Exchange 2007 to Exchange 2003 users

- Can send mail from Exchange 2007 to Internet

- Routing connectors were created for both sides during install; for my own sanity, I removed and recreated routing connectors via Exchange 2007 EMS

- Can telnet from Exchange 2003 to Exchange 2007; mail is delivered without error

- smart host is NOT configured in SMTP virtual directory

- ADSIEDIT shows correct permissions set for each administrative group (according to this link: https://www.experts-exchange.com/Software/Server_Software/Email_Servers/Exchange/Q_24557328.html?sfQueryTermInfo=1+2003+2007+destin+exchang+mail+queue+unreach)

- Ran Best Practices and Troubleshooter on both servers; both tell me message are in the queue and that routes between them 'may not be available' (obvious??)

- Windows Firewall is disable on both servers

- No Event Viewer errors on either server

- Exchange 2007 has 2 receive connectors: Client and Default. One is configured for port 587 and other is configured for port 25. Permission for port 25: anon, exchange servers, legacy exchange servers

Any ideas where to go from here?
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