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Exchange 2010 mailboxes can send mail to Exchange 2003 mailboxes, but cannot receive from them

Ok, I am so not an Exchange dude but here goes;

I've installed Exchange 2010 hub transport, client access, & mailbox roles on member server in coexistence with Exchange 2003. Exchange 2010 test mailboxes can send & receive mail from Exchange 2010 mailboxes (from local MDB) and can send mail to Exchange 2003 mailboxes and to mailboxes outside over internet, BUT Exchange 2010 mailboxes cannot receive mail from my Exchange 2003 mailboxes nor from mailboxes outside over internet. This sounds like a connector problem to me, but I cannot figure out what the dealio is. Any and all help would be ginormously appreciated.
Avatar of MegaNuk3
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Where is port 25 from the internet coming into? The exchange 2003 server?

You need an edge transport server installed or an anonymous receive connector on a hub transport to receive mail from the Internet direct to exchange 2010
During the coexistence setup a 2 way routing group connector should have been setup, has it? And are your exchange 2003 servers members of the exchange legacy interop group?
Avatar of craigerino


port 25 goes to spam filter which points to Exchange 03. I have anonymous receive from exchange 03. I do not see an exchange legacy interop group. Where would I find that?
Avatar of Jon Brelie
On your 2003 server, create a routing group connector that connects to your Ex2010 routing group.  Remote bridgehead should be your Ex2010 server.
What NDR do exchange 2003 users get when trying to mail exchange 2010 users?
Routing group that connects to Exch2010 with exchange2010 as bridgehead already present on Exchange 03. Below is NDR - EMAIL2 is Exchange 2010 & recipient "john Doe" is mailbox on Exchange 03

Generating server:
#550 5.1.1 RESOLVER.ADR.ExRecipNotFound; not found ##
Original message headers:

Received: from ([::1]) by ([::1]) with mapi id 14.01.0218.012; Wed, 10 Nov
 2010 11:56:38 -0600
Content-Type: application/ms-tnef; name="winmail.dat"
Content-Transfer-Encoding: binary
From: Email Account <>
To: John Doe
Subject: test
Thread-Topic: test
Thread-Index: AcuBAJ8YpDvgtCaiR7Kv9/T5lw8SRg==
Date: Wed, 10 Nov 2010 11:56:37 -0600
Message-ID: <>
Accept-Language: en-US
Content-Language: en-US
X-MS-TNEF-Correlator: <>
MIME-Version: 1.0
X-Originating-IP: [10.x.x.x]
Search for the exchangelegacyinterop group it should reside under the "Exchange Security Groups" OU.

Please verify that you can't email from E2k3 to E2k10?

Also confirm that under the recipient\mailbox OU under E2k10 EMC you can see mailboxes from E2k3 and E2k10
Had to rebuild the SMTP connectors
Do you mean you recreated the RGCs?
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Jon Brelie
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had to create the connectors on 2010 box, Enphyniti suggested 2003 box.
Fascinating.  I've not yet run into an issue like that.  Did you need to alter anything from the default when you recreated them, or did just recreating them do the trick?
As the issue was with the Ex2010 connectors, not 2003, I rescind my objection.