Inbound mail rejected unable to relay

I have been having some weird issues tonight, which initially presented itself as an autodiscover issue, and an expired certificate (the certificate in question had actually expired six months ago and had been replaced by a valid certificate).  I was finally able to resolve that issue by adding a registry entry to disable loopback.  I also removed the expired certificate and re-enabled the current certificate for IMAP, POP, SMTP and IIS.  After a reboot I had a conflict between the Default Web Site and SBS Web Applications, but if Default Web Site was stopped, OWA and Autodiscover work.  But now I have an even worse problem, one that I can't understand in this series of events - if I try to send an email from outside the domain to any of the users, it bounces back as Unable to Relay.  So no outside mail is getting in now and I can't figure out why.  Any help would be very much appreciated - this is extremely urgent.  Thank you!
landiiiks2Asked:
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Rajkumar-MCITPCommented:
Is that a SBS box?

All the exchange server related services are running normal? Test-ServiceHealth

Anonymous authentication is enabled on the default server name receive connector? server configuration -> Hub Transport

Did you check your domain name status, from where you bought, is that expired? MX record configuration are correctly pointed to your static IP?
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landiiiks2Author Commented:
Yes, SBS.  All services running normally.  Anonymous authentication enabled.  I did think about domain name expiration especially because it could have tied all these weird things together, but no, good till fall of '14.  MX record is fine - everything's been working perfectly till today.
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Rajkumar-MCITPCommented:
Can you paste the error message that you are receiving ? 5.7.1?

Did you check the message header of the NDR, is that your exchange server sending those NDR?

Any event logs  related to Message Transport?
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landiiiks2Author Commented:
Interesting that you asked the last question - there had not been any errors in the log all the way along, but because you answered the question I went and looked again, and this time there was.  It said "Receive Connector Requires Transport Layer Security before the MailFrom command can be run, but the server can't achieve it."  The receive connector settings were actually correct, but somehow looking all that up led me to an article that says that the SMTP service cannot be installed and running on a 2007 server.  I looked, and there it was.  I stopped it, restarted Exchange transport, and mail started going through.  I will research this more tomorrow, but there has been a database software company on that server installing what they need for this new database, and I suspect that they installed something that interfered with all of this.  Something in the article mentioned that even though the SMTP service isn't part of the Exchange installation, sometimes something else can get installed that includes it.  I have to think that's what happened, and that it also somehow interfered with the certificate.  I want to really thank you for getting back to me and ulitmately leading me to the solution.  And for allowing me to get at least a bit of sleep tonight!
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