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Migration from Exchange 2003 to 2010 leads to block of all incoming e-mail after removal of Routing Group Connector

I recently came close to completing migration from Exchange 2003 to 2010, moving from a Windows 2003 platform to Windows 2012. After I had moved all of the mailboxes everything was working fine with the systems connected. Now I need to remove Exchange 2003 from the old server. However, as soon as I execute the [Get-RoutingGroupConnector | Remove-RoutingGroupConnector] command in PowerShell on the 2010 system, all incoming e-mail is blocked somehow. Outbound email is fine. Restoring the RGC fixes the problem, but I don't want to run the old server forever. I don't see any problems with SMTP or firewalls. MX points to the new server. Please help me cut the cord to my old server before it dies on its own.
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Alan Hardisty
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In the Exchange Management Console on the Exchange 2010 server > expand Server Configuration > click on Hub Transport  > click on the name of your Exchange 2010 server in the top pane > in the Receive Connectors tab underneath right click on the Default <EX2010 Server Name> > PropertiesPermissions Group tab > make sure the Anonymous users box is ticked.
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JohnDoctor

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Thanks, but anonymous user permission was enabled in default receive connector and the problem persisted.
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VB ITS
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Did you update the port forwarding rule on your firewall as Alan suggested above to forward SMTP traffic (port 25) to the IP address of your new Exchange server?
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JohnDoctor

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Thank you, the firewall was already redirecting port 25 to the new server and we cannot get incoming email without the RGC.
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Exchange
Exchange

Exchange is the server side of a collaborative application product that is part of the Microsoft Server infrastructure. Exchange's major features include email, calendaring, contacts and tasks, support for mobile and web-based access to information, and support for data storage.

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